Read online “Human Physiology. Human physiology

Alexey Solodkov, Elena Sologub

Human physiology. General. Sports. Age

Textbook for higher educational institutions of physical culture. 7th edition

Approved by the Ministry of the Russian Federation for Physical Culture and Sports as a textbook for higher educational institutions of physical culture

The publication was prepared at the Department of Physiology of the National State University of Physical Culture, Sports and Health. P. F. Lesgafta, St. Petersburg

Reviewers:

V. I. Kuleshov, doctor med. sciences, prof. (VmedA named after S. M. Kirov)

I. M. Kozlov, Doctor of Biology and doctor ped. sciences, prof. (NSU named after P.F. Lesgaft, St. Petersburg)

* * *

Aleksey Sergeevich Solodkov – Professor of the Department of Physiology of the National State University of Physical Culture, Sports and Health named after. P. F. Lesgafta (head of the department for 25 years, 1986–2012).

Honored Scientist of the Russian Federation, Academician of the Petrovsky Academy of Sciences and Arts, Honorary Worker of Higher Professional Education of the Russian Federation, Chairman of the section “Physiology of Sports” and member of the Board of the St. Petersburg Physiological Society named after. I. M. Sechenov.

Sologub Elena Borisovna – Doctor of Biological Sciences, Professor. Since 2002 he has lived in New York (USA).

At the Department of Physiology of the National State University of Physical Culture, Sports and Health. P.F. Lesgafta worked since 1956, from 1986 to 2002 - as a professor of the department. She was elected academician of the Russian Academy of Medical and Technical Sciences, Honorary Worker of Higher Education in Russia, member of the Board of the St. Petersburg Society of Physiologists, Biochemists and Pharmacologists named after. I. M. Sechenov.

Preface

Human physiology is the theoretical basis of a number of practical disciplines (medicine, psychology, pedagogy, biomechanics, biochemistry, etc.). Without understanding the normal course of physiological processes and the constants that characterize them, various specialists cannot correctly assess the functional state of the human body and its performance in various operating conditions. Knowledge of the physiological mechanisms of regulation of various body functions is important in understanding the course of recovery processes during and after intense muscular labor.

By revealing the basic mechanisms that ensure the existence of an entire organism and its interaction with the environment, physiology makes it possible to clarify and study the conditions and nature of changes in the activity of various organs and systems in the process of human ontogenesis. Physiology is the science that carries out systems approach in the study and analysis of the diverse intra- and intersystem relationships of the complex human body and their reduction into specific functional formations and a unified theoretical picture.

It is important to emphasize that domestic researchers play a significant role in the development of modern scientific physiological concepts. Knowledge of the history of any science is a necessary prerequisite for a correct understanding of the place, role and significance of the discipline in the content of the socio-political status of society, its influence on this science, as well as the influence of science and its representatives on the development of society. Therefore, consideration of the historical path of development of individual sections of physiology, mention of its most prominent representatives and analysis of the natural science base on which the basic concepts and ideas of this discipline were formed make it possible to assess the current state of the subject and determine its further promising directions.

Physiological science in Russia in the 18th–19th centuries is represented by a galaxy of brilliant scientists - I. M. Sechenov, F. V. Ovsyannikov, A. Ya. Danilevsky, A. F. Samoilov, I. R. Tarkhanov, N. E. Vvedensky and etc. But only I.M. Sechenov and I.P. Pavlov deserve the credit for creating new directions not only in Russian, but also in world physiology.

Physiology as an independent discipline began to be taught in 1738 at the Academic (later St. Petersburg) University. Moscow University, founded in 1755, also played a significant role in the development of physiology, where the Department of Physiology was opened within its structure in 1776.

In 1798, the Medical-Surgical (Military Medical) Academy was founded in St. Petersburg, which played an exceptional role in the development of human physiology. The Department of Physiology created under her was successively headed by P. A. Zagorsky, D. M. Vellansky, N. M. Yakubovich, I. M. Sechenov, I. F. Tsion, F. V. Ovsyannikov, I. R. Tarkhanov, I. P. Pavlov, L. A. Orbeli, A. V. Lebedinsky, M.P. Brestkin and other outstanding representatives of physiological science. Behind each named name there are discoveries in physiology that are of global significance.

Physiology was included in the curriculum of physical education universities from the first days of their organization. At the Higher Courses of Physical Education created by P. F. Lesgaft in 1896, a physiology office was immediately opened, the first director of which was Academician I. R. Tarkhanov. In subsequent years, physiology was taught here by N.P. Kravkov, A.A. Walter, P.P. Rostovtsev, V.Ya. Chagovets, A. G. Ginetsinsky, A. A. Ukhtomsky, L. A. Orbeli, I. S. Beritov, A. N. Krestovnikov, G. V. Folbort and others.

The rapid development of physiology and the acceleration of scientific and technological progress in the country led to the emergence in the 30s of the 20th century of a new independent section of human physiology - sports physiology, although individual works devoted to the study of body functions during physical activity were published at the end of the 19th century (I O. Rozanov, S. S. Gruzdev, Yu. V. Blazhevich, P. K. Gorbachev, etc.). It should be emphasized that systematic research and teaching of sports physiology began in our country earlier than abroad, and was more targeted. By the way, we note that only in 1989 the General Assembly of the International Union of Physiological Sciences decided to create a commission under it “Physiology of Sports”, although similar commissions and sections in the system of the USSR Academy of Sciences, the USSR Academy of Medical Sciences, the All-Union Physiological Society named after. I. P. Pavlova of the USSR State Sports Committee have existed in our country since the 1960s.

The theoretical prerequisites for the emergence and development of sports physiology were created by the fundamental works of I. M. Sechenov, I. P. Pavlov, N. E. Vvedensky, A. A. Ukhtomsky, I. S. Beritashvili, K. M. Bykov and others. However, the systematic study of the physiological foundations of physical culture and sports began much later. Particularly great merit in the creation of this section of physiology belongs to L. A. Orbeli and his student A. N. Krestovnikov, and it is inextricably linked with the formation and development of the University of Physical Culture. P.F. Lesgaft and his department of physiology - the first such department among physical education universities in the country and in the world.

After the creation in 1919 of the Department of Physiology at the Institute of Physical Education. P. F. Lesgaft teaching this subject carried out by L. A. Orbeli, A. N. Krestovnikov, V. V. Vasilyeva, A. B. Gandelsman, E. K. Zhukov, N. V. Zimkin, A. S. Mozzhukhin, E. B. Sologub, A. S. Solodkov and others. In 1938, A. N. Krestovnikov published the first “Textbook of Physiology” in our country and in the world for physical education institutes, and in 1939 – the monograph “Physiology of Sports”. An important role in the further development of teaching the discipline was played by three editions of the “Textbook of Human Physiology” edited by N.V. Zimkin (1964, 1970, 1975).

The development of sports physiology was largely due to extensive fundamental and applied research on the subject. The development of any science poses more and more new practical problems for representatives of many specialties, to which theory cannot always and immediately provide an unambiguous answer. However, as D. Crowcroft (1970) wittily noted, “...scientific research has one strange feature: it has a habit, sooner or later, of being useful to someone or something.” Analysis of the development of educational and scientific areas of sports physiology clearly confirms this position.

The demands of the theory and practice of physical education and training require physiological science to reveal the peculiarities of the functioning of the body, taking into account the age of people and the patterns of their adaptation to muscular activity. The scientific principles of physical education of children and adolescents are based on the physiological laws of human growth and development at different stages of ontogenesis. In the process of physical education, it is necessary not only to increase motor readiness, but also to form the necessary psychophysiological properties and qualities of the individual, ensuring her readiness for work and active activity in the modern world.

The textbook was prepared in accordance with the new program in physiology for universities of physical education and the requirements of the State Standard of Higher Professional Education.
For undergraduates, graduate students, researchers, teachers, trainers and doctors working in the field of physical education.

PREFACE...... 3 Part I. GENERAL PHYSIOLOGY...... 8 1. Introduction. History of physiology...... 8 1. 1. The subject of physiology, its connection with other sciences and significance for physical culture and sports...... 8 1. 2. Methods of physiological research...... 9 1 3. Brief history of physiology...... 10 2. General principles of physiology and its basic concepts...... 12 2. 1. Basic functional characteristics of excitable tissues...... 12 2. 2. Nervous and humoral regulation of functions...... 14 2. 3. Reflex mechanism of the nervous system...... 15 2. 4. Homeostasis...... 16 2. 5. The occurrence of excitation and its implementation.. .... 17 3. Nervous system...... 21 3. 1. Basic functions of the central nervous system...... 21 3. 2. Basic functions and interactions of neurons...... 21 3. 3. Features of the activity of nerve centers...... 25 3. 4. Coordination of the activity of the central nervous system...... 29 3. 5. Functions of the spinal cord and subcortical parts of the brain...... 33 3. 6. Autonomic nervous system...... 39 3. 7. Limbic system...... 43 3. 8. Functions of the cerebral cortex...... 43 4. Higher nervous activity...... 49 4. 1. Conditions of formation and types of conditioned reflexes...... 49 4. 2. External and internal inhibition of conditioned reflexes...... 52 4. 3. Dynamic stereotype...... 52 4. 4. Types higher nervous activity, first and second signaling system...... 53 5. Neuromuscular apparatus...... 55 5. 1. Functional organization of skeletal muscles...... 55 5. 2. Mechanisms of contraction and relaxation of muscle fiber...... 57 5. 3. Single and tetanic contraction. Electromyogram...... 60 5. 4. Morphofunctional bases of muscle strength...... 63 5. 5. Modes of muscle operation...... 67 5. 6. Energy of muscle contraction...... 68 6. Voluntary movements...... 71 6. 1. Basic principles of movement organization...... 71 6. 2. The role of various parts of the central nervous system in the regulation of postural-tonic reactions...... 75 6. 3. The role of various parts of the central nervous system in the regulation of movements...... 77 6. 4. Descending motor systems...... 81 7. Sensory systems...... 83 7. 1. General plan of organization and functions sensory systems...... 83 7. 2. Classification and mechanisms of excitation of receptors...... 84 7. 3. Properties of receptors...... 86 7. 4. Coding of information...... 87 7. 5. Visual sensory system...... 88 7. 6. Auditory sensory system...... 93 7. 7. Vestibular sensory system...... 96 7. 8. Motor sensory system ...... 99 7. 9. Sensory systems of the skin, internal organs, taste and smell. ..... 102 7. 10. Processing, interaction and significance of sensory information...... 105 8. Blood...... 109 8. 1. Composition, volume and functions of blood...... 110 8. 2. Formed elements of blood...... 112 8. 3. Physico-chemical properties of blood plasma...... 116 8. 4. Blood coagulation and transfusion...... 118 8. 5 Regulation of the blood system...... 121 9. Blood circulation...... 123 9. 1. The heart and its physiological properties...... 123 9. 2. Blood movement through the vessels (hemodynamics)... .... 128 9. 3. Regulation of the cardiovascular system...... 132 10. Breathing...... 136 10. 1. External respiration...... 136 10. 2. Exchange of gases in the lungs and their transport by blood...... 139 10. 3. Regulation of breathing...... 143 11. Digestion...... 145 11. 1. General characteristics of digestive processes...... 145 11. 2. Digestion in various parts of the gastrointestinal tract...... 147 11. 3. Absorption of food digestion products...... 153 12. Metabolism and energy...... 155 12. 1. Protein metabolism...... 155 12. 2. Carbohydrate metabolism...... 156 12. 3. Lipid metabolism...... 157 12. 4. Exchange of water and mineral salts.... .. 159 12. 5. Energy exchange...... 160 12. 6. Regulation of metabolism and energy...... 163 13. Excretion...... 165 13. 1. General characteristics of excretory processes ...... 165 13. 2. Kidneys and their functions...... 165 13. 3. The process of urine formation and its regulation...... 168 13. 4. Homeostatic function of the kidneys..... 170 13. 5. Urination and urination...... 170 13. 6. Sweating...... 171 14. Heat exchange...... 173 14. 1. Human body temperature and isothermia.. .... 173 14. 2. Mechanisms of heat generation...... 174 14. 3. Mechanisms of heat transfer...... 176 14. 4. Regulation of heat exchange...... 177 15. Internal secretion.. .... 178 15. 1. General characteristics of the endocrine system...... 178 15. 2. Functions of the endocrine glands...... 181 15. 3. Changes in endocrine functions under various conditions..... 192 Part II. SPORTS PHYSIOLOGY...... 198 Section I. GENERAL SPORTS PHYSIOLOGY...... 198 1. Sports physiology - an educational and scientific discipline...... 199 1. 1. Sports physiology, its content and objectives. ..... 199 1. 2. The Department of Physiology and its role in the formation and development of sports physiology...... 201 1. 3. State and prospects for the development of sports physiology....... 206 2. Adaptation to physical loads and reserve capabilities of the body...... 210 2. 1. Dynamics of body functions during adaptation and its stages...... 211 2. 2. Physiological features of adaptation to physical activity. ..... 215 2. 3. Urgent and long-term adaptation to physical activity...... 217 2. 4. Functional adaptation system...... 221 2. 5. The concept of the physiological reserves of the body... ... 224 3. Functional states of athletes...... 226 3. 1. General characteristics of functional states...... 226 3. 2. Physiological patterns of development of functional states...... 229 3. 3 Types of functional states...... 231 4. Functional changes in the body during physical activity...... 237 4. 1. Changes in the functions of various organs and systems of the body...... 237 4. 2. Functional shifts under loads of constant power...... 240 4. 3. Functional shifts under loads of variable power...... 241 4. 4. Applied significance of functional changes for assessing the performance of athletes...... 243 5. Physiological characteristics of the body's states during sports activity...... 244 5. 1. The role of emotions during sports activity...... 244 5. 2. Pre-start states...... 247 5. 3. Warm-up and warm-up ...... 250 5. 4. Steady state during cyclic exercises...... 252 5. 5. Special states of the body during acyclic, static and variable power exercises....... 253 6. Physical performance of an athlete ...... 254 6. 1. The concept of physical performance and methodological approaches to its definition...... 255 6. 2. Principles and methods of testing physical performance...... 257 6. 3. Communication physical performance with the focus of the training process in sports...... 262 6. 4. Reserves of physical performance...... 264 7. Physiological bases of fatigue in athletes...... 269 7. 1. Definition and physiological mechanisms development of fatigue...... 269 7. 2. Factors of fatigue and the state of body functions...... 273 7. 3. Features of fatigue during various types of physical activity...... 275 7. 4. Pre-fatigue, chronic fatigue and overwork...... 278 8. Physiological characteristics of recovery processes...... 281 8. 1. General characteristics of recovery processes...... 281 8. 2. Physiological mechanisms of recovery processes... ... 283 8. 3. Physiological patterns of recovery processes...... 285 8. 4. Physiological measures to increase the efficiency of recovery...... 288 Section II. PRIVATE SPORTS PHYSIOLOGY...... 291 9. Physiological classification and characteristics of physical exercises...... 291 9. 1. Various criteria for the classification of exercises. ..... 292 9. 2. Modern classification of physical exercises...... 293 9. 3. Physiological characteristics of sports poses and static loads....... 294 9. 4. Physiological characteristics of standard cyclic and acyclic movements ...... 298 9. 5. Physiological characteristics of non-standard movements...... 303 10. Physiological mechanisms and patterns of development of physical qualities...... 305 10. 1. Forms of manifestation, mechanisms and reserves for the development of strength ...... 306 10. 2. Forms of manifestation, mechanisms and reserves for the development of speed...... 310 10. 3. Forms of manifestation, mechanisms and reserves for the development of endurance...... 313 10. 4. Concept about agility and flexibility. Mechanisms and patterns of their development...... 318 11. Physiological mechanisms and patterns of formation of motor skills...... 320 11. 1. Motor skills, skills and methods of their research...... 320 11. 2 Physiological mechanisms of the formation of motor skills...... 321 11. 3. Physiological patterns and stages of the formation of motor skills...... 324 11. 4. Physiological foundations of improving motor skills...... 330 12. Physiological basics of fitness development...... 333 12. 1. Physiological characteristics of training and fitness state...... 334 12. 2. Testing the functional readiness of athletes at rest...... 336 12. 3. Testing the functional readiness preparedness of athletes under standard and extreme loads...... 339 12. 4. Physiological characteristics of overtraining and overexertion...... 343 13. Sports performance in special environmental conditions...... 346 13. 1. The influence of temperature and air humidity on sports performance...... 346 13. 2. Sports performance under conditions of changed barometric pressure...... 348 13. 3. Sports performance under changing climatic conditions..... 353 13. 4. Physiological changes in the body during swimming...... 355 14. Physiological foundations of sports training for women...... 357 14. 1. Morphofunctional characteristics of the female body...... 357 14. 2. Changes in body functions during training...... 365 14. 3. The influence of the biological cycle on the performance of women...... 370 14. 4. Individualization of the training process, taking into account the phases of the biological cycle...... 373 15. Physiological-genetic features of sports selection...... 375 15. 1. Physiological-genetic approach to issues of sports selection...... 376 15. 2. Hereditary influences on the morpho-functional characteristics and physical qualities of a person...... 378 15. 3. Taking into account the physiological and genetic characteristics of a person in sports selection...... 383 15. 4. The meaning of genetically adequate and inadequate choice of sports activity and sensorimotor dominance...... 390 15. 5. Using genetic markers to search for highly and rapidly trained athletes...... 395 16. The influence of the genome on the functional state, performance and health of athletes.. .... 398 16. 1. Storage, transmission of hereditary information and genome decoding...... 398 16. 2. Genetic DNA markers in sports.... 402 16. 3. Genetic doping in sports.. .... 405 16. 4. Detection of doping...... 415 16. 5. Risk to health...... 417 17. Physiological foundations of health-improving physical culture...... 421 17. 1. The role of physical culture in the conditions of modern life...... 422 17. 2. Hypokinesia, physical inactivity and their influence on the human body...... 425 17. 3. The main forms of health-improving physical culture and their influence on the functional state of the body ...... 428 Part III. AGE PHYSIOLOGY...... 435 1. General physiological patterns of growth and development of the human body...... 435 1. 1. Periodization and heterochronicity of development...... 435 1. 2. Sensitive periods... ... 438 1. 3. The influence of heredity and environment on the development of the body...... 441 1. 4. Epochal and individual acceleration, biological and passport age...... 444 2. Physiological characteristics of the body of preschool children and primary school age and their adaptation to physical activity...... 448 2. 1. Development of the central nervous system, higher nervous activity and sensory systems...... 448 2. 2. Physical development and the musculoskeletal system ...... 456 2. 3. Features of blood, circulation and respiration...... 457 2. 4. Features of digestion, metabolism and energy...... 461 2. 5. Features of thermoregulation, processes secretion and activity of the endocrine glands...... 462 2. 6. Physiological features of adaptation of children of preschool and primary school age to physical activity...... 466 3. Physiological features of the body of children of middle and high school age and their adaptation to physical activity...... 488 3. 1. Development of the central nervous system, higher nervous activity and sensory systems...... 489 3. 2. Physical development and the musculoskeletal system... ... 494 3. 3. Features of blood, circulation and respiration...... 497 3. 4. Features of digestion, excretion and endocrine system...... 500 3. 5. Features of thermoregulation, metabolism and energy ...... 506 3. 6. Physiological features of adaptation of children of middle and high school age to physical activity...... 508 4. Physiological features of a physical education lesson at school...... 530 4. 1. Physiological justification for rationing physical activity for school-age children...... 530 4. 2. Changes in body functions of schoolchildren during a physical education lesson...... 533 4. 3. The influence of physical education classes on physical, functional development, performance and the state of health of schoolchildren...... 536 4. 4. Physiological and pedagogical control over physical education classes and physiological criteria for restoring the body of schoolchildren...... 543 5. Physiological characteristics of the body of mature and elderly people and their adaptation to physical activity...... 548 5. 1. Aging, life expectancy, adaptive reactions and reactivity of the body...... 549 5. 2. Age-related characteristics of the musculoskeletal system, autonomic and sensory systems..... 553 5. 3. Age-related characteristics of regulatory systems...... 557 5. 4. Physiological characteristics of adaptation of mature and elderly people to physical activity.... 561 6. Physiological characteristics of information processing in athletes of different ages. ..... 573 6. 1. The importance of information processing processes for sports and their age-related characteristics...... 573 6. 2. Physiological foundations of the processes of perception, decision-making and response programming...... 575 6 3. Speed and efficiency of tactical thinking. Brain bandwidth...... 579 6. 4. Noise immunity of athletes, its age-related characteristics...... 582 7. Functional asymmetries of athletes of different ages...... 583 7. 1. Motor asymmetries in humans, their age characteristics...... 583 7. 2. Sensory and mental asymmetries. Individual asymmetry profile...... 586 7. 3. Manifestation of functional asymmetry in athletes...... 589 7. 4. Physiological bases of training process management taking into account functional asymmetry...... 593 8. Physiological bases individual typological characteristics of athletes and their development in ontogenesis...... 595 8. 1. Individual typological characteristics of a person...... 596 8. 2. Development of typological characteristics in ontogenesis. ..... 598 8. 3. Individual typological characteristics of athletes and their consideration in the training process...... 601 8. 4. Individual typological characteristics of biorhythms and their influence on human performance...... 604 CONCLUSION...... 609

Publisher: "Sport" (2015)

Author Alexander Sergeevich Solodkov

Alexey Solodkov, Elena Sologub

Human physiology. General. Sports. Age

Textbook for higher educational institutions of physical culture

6th edition, revised and expanded

Approved by the Ministry of the Russian Federation for Physical Culture and Sports as a textbook for higher educational institutions of physical culture

The publication was prepared at the Department of Physiology of the National State University of Physical Culture, Sports and Health named after P.F. Lesgafta, St. Petersburg

Reviewers:

IN AND. Kuleshov, doctor med. sciences, prof. (VmedA named after S.M. Kirov)

THEM. Kozlov, doctor of biol, and doctor of ped. sciences, prof.

(NSU named after P.F. Lesgaft, St. Petersburg)

Preface

Human physiology is the theoretical basis of a number of practical disciplines (medicine, psychology, pedagogy, biomechanics, biochemistry, etc.) Without understanding the normal course of physiological processes and the constants that characterize them, various specialists cannot correctly assess the functional state of the human body and its performance in various conditions activities. Knowledge of the physiological mechanisms of regulation of various body functions is important in understanding the course of recovery processes during and after intense muscular labor.

By revealing the basic mechanisms that ensure the existence of an entire organism and its interaction with the environment, physiology makes it possible to clarify and study the conditions and nature of changes in the activity of various organs and systems in the process of human ontogenesis. Physiology is the science that carries out systems approach in the study and analysis of the diverse intra- and intersystem relationships of the complex human body and their reduction into specific functional formations and a unified theoretical picture.

It is important to emphasize that domestic researchers play a significant role in the development of modern scientific physiological concepts. Knowledge of the history of any science is a necessary prerequisite for a correct understanding of the place, role and significance of the discipline in the content of the socio-political status of society, its influence on this science, as well as the influence of science and its representatives on the development of society. Therefore, consideration of the historical path of development of individual sections of physiology, mention of its most prominent representatives and analysis of the natural scientific base on which the basic concepts and ideas of this discipline were formed make it possible to assess the current state of the subject and determine its further promising directions.

Physiological science in Russia in the 18th–19th centuries was represented by a galaxy of brilliant scientists - I.M. Sechenov, F.V. Ovsyannikov, A.Ya. Danilevsky, A.F. Samoilov, I.R. Tarkhanov, N.E. Vvedensky and others. But only I.M. Sechenov and I.P. Pavlov is credited with creating new directions not only in Russian, but also in world physiology.

Physiology as an independent discipline began to be taught in 1738 at the Academic (later St. Petersburg) University. Moscow University, founded in 1755, also played a significant role in the development of physiology, where the Department of Physiology was opened within its structure in 1776.

In 1798, the Medical-Surgical (Military Medical) Academy was founded in St. Petersburg, which played an exceptional role in the development of human physiology. The Department of Physiology created under her was successively headed by P.A. Zagorsky, D.M. Vellansky, N.M. Yakubovich, I.M. Sechenov, I.F. Zion, F.V. Ovsyannikov, I.R. Tarkhanov, I.P. Pavlov, L.A. Orbeli, A.V. Lebedinsky, M.P. Brestkin and other outstanding representatives of physiological science. Behind each named name there are discoveries in physiology that are of global significance.

Physiology was included in the curriculum of physical education universities from the first days of their organization. On created by P.F. Lesgaft in 1896 immediately opened a physiology cabinet at the Higher Courses of Physics Education, the first head of which was Academician I.R. Tarkhanov. In subsequent years, physiology was taught here by N.P. Kravkov, A.A. Walter, P.P. Rostovtsev, V.Ya. Chagovets, A.G. Ginetsinsky, A.A. Ukhtomsky, L.A. Orbeli, I.S. Beritov, A.N. Krestovnikov, G.V. Folbort et al.

The rapid development of physiology and the acceleration of scientific and technological progress in the country led to the emergence in the 30s of the 20th century of a new independent section of human physiology - sports physiology, although individual works devoted to the study of body functions during physical activity were published at the end of the 19th century (I O. Rozanov, S.S. Gruzdev, Yu.V. Blazhevich, P.K. Gorbachev, etc.). It should be emphasized that systematic research and teaching of sports physiology began in our country earlier than abroad, and was more targeted. By the way, we note that only in 1989 the General Assembly of the International Union of Physiological Sciences decided to create a commission under it “Physiology of Sports”, although similar commissions and sections in the system of the USSR Academy of Sciences, the USSR Academy of Medical Sciences, the All-Union Physiological Society named after. I.P. Pavlov State Sports Committee of the USSR existed in our country since the 1960s.

The theoretical prerequisites for the emergence and development of sports physiology were created by the fundamental works of I.M. Sechenova, I.P. Pavlova, N.E. Vvedensky, A.A. Ukhtomsky, I.S. Beritashvili, K.M. Bykov and others. However, the systematic study of the physiological foundations of physical culture and sports began much later. Especially great credit for the creation of this section of physiology belongs to L.A. Orbeli and his student A.N. Krestovnikov, and it is inextricably linked with the formation and development of the University of Physical Culture named after. P.F. Lesgaft and its Department of Physiology - the first such department among physical education universities in the country and in the world.

After the creation in 1919 of the Department of Physiology at the Institute of Physical Education. P.F. Lesgaft, this subject was taught by L.A. Orbeli, A.N. Krestovnikov, V.V. Vasilyeva, A.B. Gandelsman, E.K. Zhukov, N.V. Zimkin, A.S. Mozzhukhin, E.B. Sologub, A.S. Solodkov and others. In 1938 A.N. Kreetovnikov published the first “Textbook of Physiology” in our country and in the world for physical education institutes, and in 1939, the monograph “Physiology of Sports”. An important role in the further development of teaching the discipline was played by three editions of the “Textbook of Human Physiology” edited by N.V. Zimkina (1964, 1970, 1975).

The formation of various organs and systems, motor qualities and skills, their improvement in the process of physical education can be successful subject to the scientifically based use of various means and methods of physical culture, as well as if it is necessary to intensify or reduce muscle loads. In this case, it is necessary to take into account the age-sex and individual characteristics of children, adolescents, mature and elderly people, as well as the reserve capabilities of their body at different stages of individual development. Knowledge of such patterns by specialists will protect the practice of physical education from the use of both insufficient and excessive muscle loads that are dangerous to people’s health.

To date, significant factual materials on sports and age-related physiology have been accumulated, presented in relevant textbooks and teaching aids. However, in recent years, new data has appeared on some sections of the subject that were not included in previous publications. In addition, due to the constantly changing and supplemented curriculum, the content of previously published sections of the discipline does not correspond to modern thematic plans according to which teaching is conducted in physical education universities in Russia. Taking into account the above, the proposed textbook contains systematized, supplemented and, in some cases, new materials within the framework of today's educational and scientific information on the subject. The corresponding sections of the textbook also include the results of the authors’ own research.

In 1998–2000 A.S. Solodkov and E.B. Sologub published three textbooks on general, sports and developmental physiology, which were widely in demand by students, approved by teachers and served as the basis for the preparation of a modern textbook. The textbook they published in 2001 corresponds to the new program for the discipline, the requirements of the State Standard of Higher Professional Education of the Russian Federation and includes three parts - general, sports and age physiology.

Despite the large circulation of the first edition (10 thousand copies), two years later the textbook was not available in stores. Therefore, after making some corrections and additions, in 2005 the textbook was republished in the same edition. However, by the end of 2007 it turned out to be impossible to purchase it anywhere. At the same time, the Department of Physiology regularly receives proposals from various regions of the Russian Federation and CIS countries about the need for the next re-edition of the textbook. In addition, the authors have at their disposal some new materials that meet the requirements of the Bologna Process for specialists in physical education and sports.

The prepared third edition of the textbook, along with taking into account and implementing individual comments and suggestions from readers, also includes two new chapters: “Functional state of athletes” and “The influence of the genome on the functional state, performance and health of athletes.” For the last chapter, some materials were presented by N.M., professor of the Department of Biology at St. John's University in New York. Konevoy-Hanson, for which the authors are sincerely grateful to Natalya Mikhailovna.

All comments and suggestions regarding the fifth edition, aimed at improving the quality of the textbook, will be gratefully accepted by the authors.

Part I

General physiology

For successful professional activities, any trainer and teacher needs knowledge of the functions of the human body. Only taking into account the peculiarities of its vital activity can help to properly manage the growth and development of the human body, preserve the health of children and adults, maintain performance even in old age, and rationally use muscle loads in the process of physical education and sports training.

1. Introduction. History of physiology

The date of formation of modern physiology is 1628, when the English physician and physiologist William Harvey published the results of his research on blood circulation in animals.

Physiology the science of the functions and mechanisms of activity of cells, tissues, organs, systems and the entire organism as a whole. A physiological function is the manifestation of the vital activity of the organism, which has adaptive significance.

1.1. The subject of physiology, its connection with other sciences and its significance for physical culture and sports

Physiology as a science is inextricably linked with other disciplines. It is based on knowledge of physics, biophysics and biomechanics, chemistry and biochemistry, general biology, genetics, histology, cybernetics, anatomy. In turn, physiology is the basis of medicine, psychology, pedagogy, sociology, theory and methods of physical education. In the process of development of physiological science from general physiology various private sections: labor physiology, physiology...

Textbook for higher educational institutions of physical culture. 7th edition

Approved by the Ministry of the Russian Federation for Physical Culture and Sports as a textbook for higher educational institutions of physical culture

The publication was prepared at the Department of Physiology of the National State University of Physical Culture, Sports and Health. P. F. Lesgafta, St. Petersburg

Reviewers:

V. I. Kuleshov, doctor med. sciences, prof. (VmedA named after S. M. Kirov)

I. M. Kozlov, Doctor of Biology and doctor ped. sciences, prof. (NSU named after P.F. Lesgaft, St. Petersburg)

Sologub Elena Borisovna – Doctor of Biological Sciences, Professor. Since 2002 he has lived in New York (USA).

Preface

Publisher: "Sport" (2015)